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HLA-DRB

HLA-DRB References

Allele Name Local Name Cell ID Accession Numbers Authors Title Journal Year Vol. Pages Contact
DRB1*010101 - LG2 HLA00664 Bell JI, Estess P, St John T, Saiki R, Watling DL, Erlich HA, McDevitt HO DNA sequence and characterization of human class II major histocompatibility complex beta chains from the DR1 haplotype Proc Natl Acad Sci U S A 1985 82 3405-9 Prof John Bell
DRB1*010101 - 45.1 HLA00664 Tonnelle C, DeMars R, Long EO DO beta: a new beta chain gene in HLA-D with a distinct regulation of expression EMBO J 1985 4 2839-47 Prof Eric Long
DRB1*010101 - JSA
DRH
CHG		 HLA00664 - - - - - - Prof Antonio Arnaiz-Villena
DRB1*010102 - 9380965 HLA01598 - - - - - - Dr Jin Wu
DRB1*010103 - OLLI HLA01869 Dubois V, Favre-Victoire I, Gebuhrer L Three new DRB alleles routinely identified by sequence-based typing: DRB1*010103, DRB1*0326 and DRB3*0219 Tissue Antigens 2004 64 621-3 Dr Lucette Gebuhrer
DRB1*010104 - OT11400 HLA03225 - - - - - - Dr Sofia Tavoularis
DRB1*010105 - CTM-5099261 HLA03373 - - - - - - Dr Antonio Balas
DRB1*010106 - 36573 HLA03389 - - - - - - Dr Juergen Enczmann
DRB1*010107 - 47995 HLA03390 - - - - - - Dr Juergen Enczmann
DRB1*010201 - NASC HLA00665 Merryman P, Gregersen PK, Lee S, Silver J, Nunez-Roldan A, Crapper R, Winchester R Nucleotide sequence of a DRw10 beta chain cDNA clone. Identity of the third D region with that of the DRw53 allele of the beta 2 locus and as the probable site encoding a polymorphic MHC class II epitope J Immunol 1988 140 2447-52 Prof Robert Winchester
DRB1*010201 1568C69 1568 HLA00665 Hurley CK, Ziff BL, Silver J, Gregersen PK, Hartzman R, Johnson AH Polymorphism of the HLA-DR1 haplotype in American blacks. Identification of a DR1 beta-chain determinant recognized in the mixed lymphocyte reaction J Immunol 1988 140 4019-23 Dr Carolyn K Hurley
DRB1*010201 - MUM HLA00665 - - - - - - Prof Antonio Arnaiz-Villena
DRB1*010202 DRB1*01DMT TO0973 HLA00666 Hashemi-Tavoularis S, Couture C, Buyse IM Identification of new DRB1*01 (DRB1*01022), DRB1*14 (DRB1*1428) and DRB3* (DRB3*0206) alleles Tissue Antigens 1997 50 89-93 Dr Sofia Tavoularis
DRB1*010203 - 376P01 HLA01875 Bontadini A, Manfroi S, Delfino L, Fruet F, Iannelli S, Parodi AM, Ferrara GB, Conte R Characterization of a new HLA-DRB1*01 allele (HLA-DRB1*010203) in a Caucasian Italian family by using sequence-based typing Tissue Antigens 2004 64 624-6 Dr Roberto Conte
DRB1*010204 - 84308 HLA02001 - - - - - - Dr Mary Sue Leffell
DRB1*0103 DR"Br"B1 TER-ND HLA00667 Bidwell JL, Bidwell EA, Sansom DM, Klouda PT, Bradley BA The origin of HLA-DR"Br": exon 2 nucleotide sequence implicates possible gene conversion of DR1 by DR4-Dw10, DR5, or DRw6-Dw18 Hum Immunol 1989 26 191-7 Dr Jeff Bidwell
DRB1*0103 DRB1*BON BON HLA00667 Coppin HL, Avoustin P, Fabron J, Huchenq A, Garnier JM, Thomsen M, De Preval C Evolution of the HLA-DR1 gene family. Structural and functional analysis of the new allele "DR-BON" J Immunol 1990 144 984-9 Dr Hélène Coppin
DRB1*0103 DR1-CETUS BG HLA00667 Gregersen PK, Todd JA, Erlich HA, Long E, Servenius B, Choi E, Kao HT, Lee JS First domain sequence diversity of DR and DQ subregion alleles Immunobiology of HLA. Vol. 1 1989 - 1027-31 Dr Henry Erlich
DRB1*0104 DRB1*01new LAUTH J HLA00668 Guignier F, Mercier B, Roz P, Ferec C, Saleun JP, Chatelain P A novel HLA-DRB1*01 allele (DRB1*0104) Tissue Antigens 1993 42 42-4 Dr Bernard Mercier
DRB1*0104 - LAUTH J HLA00668 - - - - - - Prof Derek Middleton
DRB1*0105 DRB1*0101V1 JC10218 HLA00669 - - - - - - Dr Kouichi Kashiwase
DRB1*0106 DRB1*01New MGM14106 HLA00670 Palou E, Mongay L, Arias MT, Isart F, Suarez B, Masso M, Fabregat V, Martorell J, Gaya A Identification of a novel DRB1-allele (DRB1*0106) by sequence-based typing Tissue Antigens 1999 53 308-10 Dr Antonio Gaya
DRB1*0107 DRB1*New ZAE HLA01307 Voorter CE, Hepkema BG, Lems SP, van den Berg-Loonen EM Identification of three new DRB1 alleles, DRB1*0107, *0425 and *13012 and confirmation of DRB4*01033 Tissue Antigens 2003 61 398-402 Prof Ella M van den Berg Loonen
DRB1*0107 - IOL Gae
IOL Ire		 HLA01307 - - - - - - Dr Anne Dormoy
DRB1*0108 DR1-BCN HSP934010 HLA01483 Martinez-Gallo M, Martinez-Lostao L, Martinez-Carretero MA, Gago I, Maroto P, Rodriguez-Sanchez JL, de la Calle-Martin O Identification of a novel HLA-DRB1 allele, DRB1*0108, by sequence-based DRB typing in two siblings Tissue Antigens 2002 59 350-1 Dr Oscar de la Calle-Martin
DRB1*0109 - G044601003890Z HLA01633 - - - - - - Dr Deborah Sage
DRB1*0110 DRB1*01CBRL CBRL9-58-288 HLA01671 - - - - - - Dr Dong-Feng Chen
DRB1*0111 - 03-2215 HLA01811 - - - - - - Dr Mats Bengtsson
DRB1*0112 - MHH0401996 HLA02056 Horn PA, DeLuca DS, Mueller K, Blasczyk R Peptide-binding characteristics of the novel allele DRB1*0112 are probably identical to DRB1*0101 Tissue Antigens 2005 65 505-6 Prof Rainer Blasczyk
DRB1*0113 - CTM-1096976 HLA02173 Balas A, Vilches C, Rodriguez MA, Fernandez B, Martinez MP, de Pablo R, Garcia-Sanchez F, Vicario JL Group-specific amplification of cDNA from DRB1 genes. Complete coding sequences of partially defined alleles and identification of the new alleles DRB1*040602, DRB1*111102, DRB1*080103, and DRB1*0113. Human Immunology 2006 67 1008-16 Dr Antonio Balas
DRB1*0114 - Xian8576 HLA02494 Liu ML. Qi J, Zhang Y, Ye SH Identification of a novel allele HLA-DRB1*0114 in two Chinese marrow donors by sequence-based typing. Tissue Antigens 2007 69 622 Dr Mengli Liu
DRB1*0115 - VTIS89969 HLA02500 - - - - - - Dr Brian D Tait
DRB1*0116 - 2006-038-07126 HLA02666 - - - - - - Mr Brian Norvell
DRB1*0116 - NT00740 HLA02666 - - - - - - Dr Carolyn K Hurley
DRB1*0117 - BY00214 HLA03037 - - - - - - Dr Carolyn K Hurley
DRB1*0118 - BY00212 HLA03038 - - - - - - Dr Carolyn K Hurley
DRB1*0119 - BY00209
BY00208		 HLA03040 - - - - - - Dr Carolyn K Hurley
DRB1*0120 - JMDP36K017 HLA03377 - - - - - - Dr Kenji Tadokoro
DRB1*030101 - WT49 HLA00671 Bell JI, Denney D Jr, Foster L, Belt T, Todd JA, McDevitt HO Allelic variation in the DR subregion of the human major histocompatibility complex Proc Natl Acad Sci U S A 1987 84 6234-8 Prof John Bell
DRB1*030101 DR3 I AVL HLA00671 Gorski J, Mach B Polymorphism of human Ia antigens: gene conversion between two DR beta loci results in a new HLA-D/DR specificity Nature 1986 322 67-70 Dr Jack Gorski
DRB1*030101 - RAJI HLA00671 Gustafsson K, Wiman K, Emmoth E, Larhammar D, Bohme J, Hyldig-Nielsen JJ, Ronne H, Peterson PA, Rask L Mutations and selection in the generation of class II histocompatibility antigen polymorphism EMBO J 1984 3 1655-61 Dr Lars Rask
DRB1*030101 - DM24
DM28
DM29		 HLA00671 Todd JA, Bell JI, McDevitt HO HLA-DQ beta gene contributes to susceptibility and resistance to insulin-dependent diabetes mellitus Nature 1987 329 599-604 Dr John A Todd
DRB1*030101 (DRB3) CMCC HLA00671 Horn GT, Bugawan TL, Long CM, Manos MM, Erlich HA Sequence analysis of HLA class II genes from insulin-dependent diabetic individuals Hum Immunol 1988 21 249-63 Dr Henry Erlich
DRB1*030101 dJ93N13 HSF7 HLA00671 - - - - - - Prof Stephan Beck
DRB1*030101 - APR
ALL
MVJ
MUR		 HLA00671 - - - - - - Prof Antonio Arnaiz-Villena
DRB1*030101 - U-STH HLA00671 Eberle M, Asu U, Taylor M, Hunter JB, Fuller TC, Maurer D Identification of a putative DR8 founder haplotype containing a novel DRB1*08 allele Hum Immunol 1994 40(S1) 77 Dr Thomas C Fuller
DRB1*030101 - QBL HLA00671 Traherne JA, Horton R, Roberts AN, Miretti MM, Hurles ME, Stewart CA, Ashurst JL, Atrazhev AM, Coggill P, Palmer S, Almeida J, Sims S, Wilming LG, Rogers J, de Jong PJ, Carrington M, Elliott JF, Sawcer S, Todd JA, Trowsdale J, Beck S Genetic analysis of completely sequenced disease-associated MHC haplotypes identifies shuffling of segments in recent human history PLoS Genetics 2006 2 81-92 Prof Stephan Beck
DRB1*030102 - 21 HLA00672 Eberle M, Baxter-Lowe LA A silent mutation in HLA-DRB1*0301 can affect oligotyping Tissue Antigens 1992 40 150-2 Prof Lee Ann Baxter-Lowe
DRB1*030102 DRB1*IMR M.R. HLA00672 Buyse I, Emonds MP, Bouillon R, Marynen P, Cassiman JJ Novel class II HLA-DRB4 and DPB1 alleles found in the Belgian population Immunogenetics 1993 38 380 Prof Jean-Jaques Cassiman
DRB1*030102 - CTM-2095057 HLA00672 Balas A, Vilches C, Rodriguez MA, Fernandez B, Martinez MP, de Pablo R, Garcia-Sanchez F, Vicario JL Group-specific amplification of cDNA from DRB1 genes. Complete coding sequences of partially defined alleles and identification of the new alleles DRB1*040602, DRB1*111102, DRB1*080103, and DRB1*0113. Human Immunology 2006 67 1008-16 Dr Antonio Balas
DRB1*030103 - D22559 HLA02769 - - - - - - Miss Odile Avinens
DRB1*030104 - VTIS65942 HLA02830 - - - - - - Dr Brian D Tait
DRB1*030105 - NT00735 HLA02955 Lazaro AM, Xiao Y, Masaberg C, Lebeck L, Ng J, Hurley CK, Posch P Novel alleles at the HLA-DRB1 and -DQB1 loci. Tissue Antigens 2008 72 72-74 Dr Carolyn K Hurley
DRB1*030106 - MHHN-593833 HLA03061 - - - - - - Prof Rainer Blasczyk
DRB1*030201 - 2041
1563		 HLA00673 Hurley CK, Gregersen PK, Gorski J, Steiner N, Robbins FM, Hartzman R, Johnson AH, Silver J The DR3(w18),DQw4 haplotype differs from DR3(w17),DQw2 haplotypes at multiple class II loci Hum Immunol 1989 25 37-50 Dr Carolyn K Hurley
DRB1*030201 - 24A1 HLA00673 - - - - - - Prof Antonio Arnaiz-Villena
DRB1*030202 - GN055 HLA00674 Ellis JM, Steiner N, Wang J, Tang TT, Hurley CK Diversity and evolution of the DRB1*03 family: description of DRB1*03022,*0307,*0308 Tissue Antigens 1997 49 41-5 Dr Carolyn K Hurley
DRB1*030202 - GMONT (ES) HLA00674 - - - - - - Dr Michelle Taylor
DRB1*0303 - RBL B25 HLA00675 Apple RJ, Erlich HA Two new HLA DRB1 alleles found in African Americans: implications for balancing selection at positions 57 and 86 Tissue Antigens 1992 40 69-74 Dr Henry Erlich
DRB1*0303 - GN00265 HLA00675 - - - - - - Dr Carolyn K Hurley
DRB1*0304 MIT-3758 MIT-3758
GRD		 HLA00676 Anholts JD, Verduyn W, Parlevliet A, Doxiadis II, D'Amaro J, Giphart MJ, Persijn GG, Schreuder GM Irregular polymerase chain reaction-sequence-specific oligonucleotide hybridization patterns reveal seven new HLA-DRB1 alleles related to DR2, DR3, DR6, DR8, and DR11. Implications for sequence-specific priming Hum Immunol 1995 42 15-22 Dr Willem Verduyn
DRB1*0304 DRB1*0304 35919 HLA00676 - - - - - - Dr Martin Guttridge
DRB1*0304 - 35919
TER336		 HLA00676 - - - - - - Prof Rainer Blasczyk
DRB1*030501 DR3New U-HFI (#662) HLA00677 Asu UM, Taylor M, Dunn D, Fuller TC Identification of a new HLA-DR3 allele: DRB1*0305 Tissue Antigens 1995 46 405-7 Dr Thomas C Fuller
DRB1*030501 - TTO5607 HLA00677 - - - - - - Dr Sue Bowman
DRB1*030502 DRB1*LAHRE LAHRE HLA01428 Greville WD, Chapman G, Hogbin JP, Kennedy A, Dunckley H Novel HLA-DRB1 alleles discovered during routine sequencing based typing, DRB1*03052, DRB1*04032, DRB1*1139 and DRB1*1346 Tissue Antigens 2002 59 154-6 Dr Heather Dunckley
DRB1*0306 JV 1094 JV1094 HLA00678 Fenske B, Lemieux J, Hoar D, Fitzpatrick J, Keown P A novel DR beta 1*03 allele identified by polymerase chain reaction-restriction fragment length polymorphism Hum Immunol 1996 46 55-7 Dr Bruce Fenske
DRB1*0306 - CTM-2096757 HLA00678 Balas A, Vilches C, Rodriguez MA, Fernandez B, Martinez MP, de Pablo R, Garcia-Sanchez F, Vicario JL Group-specific amplification of cDNA from DRB1 genes. Complete coding sequences of partially defined alleles and identification of the new alleles DRB1*040602, DRB1*111102, DRB1*080103, and DRB1*0113. Human Immunology 2006 67 1008-16 Dr Antonio Balas
DRB1*0307 DRB1*03011New GN073 HLA00679 Ellis JM, Steiner N, Wang J, Tang TT, Hurley CK Diversity and evolution of the DRB1*03 family: description of DRB1*03022,*0307,*0308 Tissue Antigens 1997 49 41-5 Dr Carolyn K Hurley
DRB1*0307 - NT00982 HLA00679 - - - - - - Dr Carolyn K Hurley
DRB1*0308 DRB1*03Variant GN090

 HLA00680 Ellis JM, Steiner N, Wang J, Tang TT, Hurley CK Diversity and evolution of the DRB1*03 family: description of DRB1*03022,*0307,*0308 Tissue Antigens 1997 49 41-5 Dr Carolyn K Hurley
DRB1*0309 - D438 HLA00681 Dufosse F, Guignier F, Cracco P, Becuwe D, Huart JJ A novel HLA DRB1 allele (DRB1*0309) Tissue Antigens 1997 49 523-5 Dr Françoise Dufosse
DRB1*0310 - PMR HLA00682 Martinez-Quiles N, Martin-Villa JM, Martinez-Laso J, Perez-Blas M, Ferre-Lopez S, Moreno-Pelayo MA, Alvarez-Tejado M, Arnaiz-Villena A Description of two new HLA-DRB alleles (DRB1*0310 and DRB3*01012) found in a Spanish infant Tissue Antigens 1997 49 658-61 Prof Antonio Arnaiz-Villena
DRB1*0311 DRB1*0301Variant UWE02 HLA00683 - - - - - - Dr Carolyn K Hurley
DRB1*0312 DRB1*03AGC WVN HLA00684 Anholts JD, Verduijn W, Drabbels J, Mulder A, Doxiadis II, Schreuder GM Identification of two new alleles HLA-DRB1*0312, DRB1*0432 and of a DRB3-negative DRB1*1313-positive haplotype Tissue Antigens 2000 56 87-9 Dr Willem Verduyn
DRB1*0313 DRB1*03New DELAT HLA01007 - - - - - - Dr Anne Dormoy
DRB1*0314 DR'KW' KW HLA01008 Koninkx HJ, Tijssen HJ, Joosten I Exon 2 sequence analysis of a novel HLA-DRB1 allele, DRB1*0314 Tissue Antigens 2001 58 205-6 Dr Irma Joosten
DRB1*0315 DRB1*0301A DKMS 585607 HLA01087 - - - - - - Dr Claudia MŸller
DRB1*0315 - TER317 HLA01087 - - - - - - Prof Rainer Blasczyk
DRB1*0316 - 09343336 HLA01152 - - - - - - Dr Carolyn K Hurley
DRB1*0317 DRB1*13KM SMS202-147-KerHut HLA01153 Buhlmann A, Albis-Camps M, Holtkamp B, Blasczyk R Two novel DRB alleles detected by PCR-SSO and confirmed by sequencing: DRB1*0317 and DRB3*0210 Tissue Antigens 2000 56 553-5 Prof Rainer Blasczyk
DRB1*0318 DRB1*03XX RSA036575
MSA058812		 HLA01349 Ercilla MG, Guardia A, Suarez B, Arias MT, Fabregat V, Costa M, Salinas I, Martorell J, Vives J, Lozano F Identification of a new HLA-DRB1 allele (DRB1*0318) in three members of a Caucasian Spanish family Tissue Antigens 2001 57 489-91 Dr Francisco Lozano
DRB1*0319 DRB1*GCASS GCASS HLA01432 - - - - - - Dr Heather Dunckley
DRB1*0319 - 133983 HLA01432 - - - - - - Dr Heather Dunckley
DRB1*0320 DRB1*03011var NT0022 HLA01455 - - - - - - Dr Carolyn K Hurley
DRB1*0321 - Patient#17839 HLA01501 Gagne K, Bonneville F, Cheneau ML, Herry P, Pinson MJ, Herfray P, Cesbron-Gautier A, Follea G, Bignon JD Identification of a new HLA-DRB1 allele, DRB1*0321 Tissue Antigens 2002 60 268-70 Dr Katia Gagne
DRB1*0322 - MAWE0816AN HLA01557 - - - - - - Dr Ann-Margaret Little
DRB1*0322 - LB64975
LB62448		 HLA01557 Anholts JD, Drabbels J, Verduijn W, Oudshoorn M, Schreuder GM Five newly identified HLA alleles: A*0310, A*2907, B*4435, Cw*0206, Cw*0506, and confirmation of A*3106, B*3924, Cw*0314, DRB1*0322, and DRB1*1433 alleles Tissue Antigens 2004 63 80-4 Dr Ieke Schreuder
DRB1*0323 - DNA6060 HLA01614 - - - - - - Dr Angelika Reil
DRB1*0324 - ULM13352 HLA01687 - - - - - - Dr Kaimo Hirv
DRB1*0325 - DKM544293 HLA01692 - - - - - - Dr Kaimo Hirv
DRB1*0326 - BELF HLA01868 Dubois V, Favre-Victoire I, Gebuhrer L Three new DRB alleles routinely identified by sequence-based typing: DRB1*010103, DRB1*0326 and DRB3*0219 Tissue Antigens 2004 64 621-3 Dr Lucette Gebuhrer
DRB1*0327 - 5994 HLA01930 - - - - - - Dr Monique Bois
DRB1*0327 - BY00119 HLA01930 - - - - - - Dr Carolyn K Hurley
DRB1*0328 - NT00510 HLA01933 - - - - - - Dr Carolyn K Hurley
DRB1*0329 - NT00626 HLA02420 - - - - - - Dr Carolyn K Hurley
DRB1*0329 - BY00247 HLA02420 - - - - - - Dr Carolyn K Hurley
DRB1*0330 DRB1*03MSC0506 MHHZ-00020533 HLA02603 - - - - - - Prof Rainer Blasczyk
DRB1*0331 - TZDR03 HLA02606 Yang KL, Chu CC, Huang CH, Lin SZ, Hsu WL, Liu DW, Lin PY Discovery of HLA-DRB1*0331 in a Taiwanese marrow donor and the importance of sequence-based typing in a rare or previously unrecognized allele. Int J Immunogenetics 2007 34 91-5 Mr Edward KL Yang
DRB1*0332 - VTIS103774 HLA02827 - - - - - - Dr Brian D Tait
DRB1*0333 - VTIS200629
VTIS200663
VTIS200630
VTIS200631		 HLA02870 - - - - - - Dr Brian D Tait
DRB1*0334 - OT14365 HLA02886 - - - - - - Dr Sofia Tavoularis
DRB1*0335 - 034162 HLA02902 - - - - - - Dr Jianping Li
DRB1*0336 - LUMC-DRB15 HLA02996 - - - - - - Miss Jacqy DH Anholts
DRB1*0337 - BY00232 HLA03059 - - - - - - Dr Carolyn K Hurley
DRB1*0337 - BY00254 HLA03059 - - - - - - Dr Carolyn K Hurley
DRB1*0338 - BY00237 HLA03067 - - - - - - Dr Carolyn K Hurley
DRB1*0339 - BY00253 HLA03075 - - - - - - Dr Carolyn K Hurley
DRB1*0340 - 60648330 HLA03370 - - - - - - Dr Reinhard Kelsch
DRB1*040101 Dw4 WT51 HLA00685 Gregersen PK, Shen M, Song QL, Merryman P, Degar S, Seki T, Maccari J, Goldberg D, Murphy H, Schwenzer J, Wang CY, Winchester RJ, Nepom GT, Silver J Molecular diversity of HLA-DR4 haplotypes Proc Natl Acad Sci U S A 1986 83 2642-6 Dr Peter Gregersen
DRB1*040101 - PRIESS HLA00685 Spies T, Sorrentino R, Boss JM, Okada K, Strominger JL Structural organization of the DR subregion of the human major histocompatibility complex Proc Natl Acad Sci U S A 1985 82 5165-9 Prof Jack L Strominger
DRB1*040101 - WT51 HLA00685 Bell JI, Denney D Jr, Foster L, Belt T, Todd JA, McDevitt HO Allelic variation in the DR subregion of the human major histocompatibility complex Proc Natl Acad Sci U S A 1987 84 6234-8 Prof John Bell
DRB1*040101 MJ8.2 MJ4 HLA00685 Curtsinger JM, Hilden JM, Cairns JS, Bach FH Evolutionary and genetic implications of sequence variation in two nonallelic HLA-DR beta-chain cDNA sequences Proc Natl Acad Sci U S A 1987 84 209-13 Prof Fritz H Bach
DRB1*040101 - BOLETH HLA00685 Andersson G, Larhammar D, Widmark E, Servenius B, Peterson PA, Rask L Class II genes of the human major histocompatibility complex. Organization and evolutionary relationship of the DR beta genes J Biol Chem 1987 262 8748-58 Dr Göran Andersson
DRB1*040101 - LTC HLA00685 - - - - - - Prof Antonio Arnaiz-Villena
DRB1*040102 DR401v MC HLA00686 Thonnard J, Gervais T, Heusterspreute M, Mersch G, De Canck I, De Greef C, Demanet C, Van Waeyenberge C A new silent mutation at codon 35 in exon 2 yielding DRB1*04012 allele Tissue Antigens 1997 49 274-6 Dr Joelle Thonnard
DRB1*040103 - BY00238 HLA03066 - - - - - - Dr Carolyn K Hurley
DRB1*0402 Dw10 FS HLA00687 Gregersen PK, Shen M, Song QL, Merryman P, Degar S, Seki T, Maccari J, Goldberg D, Murphy H, Schwenzer J, Wang CY, Winchester RJ, Nepom GT, Silver J Molecular diversity of HLA-DR4 haplotypes Proc Natl Acad Sci U S A 1986 83 2642-6 Dr Peter Gregersen
DRB1*0402 - DM24 HLA00687 Todd JA, Bell JI, McDevitt HO HLA-DQ beta gene contributes to susceptibility and resistance to insulin-dependent diabetes mellitus Nature 1987 329 599-604 Dr John A Todd
DRB1*0402 (DRB4) MMCC HLA00687 Horn GT, Bugawan TL, Long CM, Manos MM, Erlich HA Sequence analysis of HLA class II genes from insulin-dependent diabetic individuals Hum Immunol 1988 21 249-63 Dr Henry Erlich
DRB1*0402 - LPB HLA00687 - - - - - - Prof Antonio Arnaiz-Villena
DRB1*0402 - YAR HLA00687 - - - - - - Dr Alies Snijders
DRB1*0402 - FS HLA00687 Ramon D, Corell A, Cox ST, Soteriou B, Madrigal JA, Marsh SG Complete cDNA sequences of the HLA-DRB1*0402 and DRB1*11041 alleles Eur J Immunogenet 2002 29 453-5 Dr Steven GE Marsh
DRB1*040301 S3.4 SSTO HLA00688 Cairns JS, Curtsinger JM, Dahl CA, Freeman S, Alter BJ, Bach FH Sequence polymorphism of HLA DR beta 1 alleles relating to T-cell-recognized determinants Nature 1985 317 166-8 Prof Fritz H Bach
DRB1*040301 - TAS HLA00688 Gregersen PK, Goyert SM, Song QL, Silver J Microheterogeneity of HLA-DR4 haplotypes: DNA sequence analysis of LD"KT2" and LD"TAS" haplotypes Hum Immunol 1987 19 287-92 Dr Peter Gregersen
DRB1*040301 - NBP HLA00688 - - - - - - Prof Antonio Arnaiz-Villena
DRB1*040302 DRB1*SD BM1 116040 HLA01009 Greville WD, Chapman G, Hogbin JP, Kennedy A, Dunckley H Novel HLA-DRB1 alleles discovered during routine sequencing based typing, DRB1*03052, DRB1*04032, DRB1*1139 and DRB1*1346 Tissue Antigens 2002 59 154-6 Dr Warwick Greville
DRB1*040302 - 32891 HLA01009 - - - - - - Dr Martin Guttridge
DRB1*040302 - TER337 HLA01009 - - - - - - Prof Rainer Blasczyk
DRB1*040303 - 173171 HLA02717 - - - - - - Dr Heather Dunckley
DRB1*040304 - NT00760 HLA03172 Lazaro AM, Xiao Y, Masaberg C, Lebeck L, Ng J, Hurley CK, Posch P Novel alleles at the HLA-DRB1 and -DQB1 loci. Tissue Antigens 2008 72 72-74 Dr Carolyn K Hurley
DRB1*0404 LS5.8.1 LS40 HLA00689 Cairns JS, Curtsinger JM, Dahl CA, Freeman S, Alter BJ, Bach FH Sequence polymorphism of HLA DR beta 1 alleles relating to T-cell-recognized determinants Nature 1985 317 166-8 Prof Fritz H Bach
DRB1*0404 Dw14 BIN40 HLA00689 Gregersen PK, Shen M, Song QL, Merryman P, Degar S, Seki T, Maccari J, Goldberg D, Murphy H, Schwenzer J, Wang CY, Winchester RJ, Nepom GT, Silver J Molecular diversity of HLA-DR4 haplotypes Proc Natl Acad Sci U S A 1986 83 2642-6 Dr Peter Gregersen
DRB1*0404 - DM29 HLA00689 Todd JA, Bell JI, McDevitt HO HLA-DQ beta gene contributes to susceptibility and resistance to insulin-dependent diabetes mellitus Nature 1987 329 599-604 Dr John A Todd
DRB1*0404 - RGR HLA00689 - - - - - - Prof Antonio Arnaiz-Villena
DRB1*040501 Dw15 LKT3 HLA00690 Gregersen PK, Shen M, Song QL, Merryman P, Degar S, Seki T, Maccari J, Goldberg D, Murphy H, Schwenzer J, Wang CY, Winchester RJ, Nepom GT, Silver J Molecular diversity of HLA-DR4 haplotypes Proc Natl Acad Sci U S A 1986 83 2642-6 Dr Peter Gregersen
DRB1*040501 - JML
AHC
CRP		 HLA00690 Morales P, Martinez-Laso J, Martin-Villa JM, Corell A, Vicario JL, Varela P, Perez-Aciego P, Arnaiz-Villena A High frequency of the HLA-DRB1*0405-(Dw15)-DQw8 haplotype in Spaniards and its relationship to diabetes susceptibility Hum Immunol 1991 32 170-5 Dr Alfredo Corell
DRB1*040501 - DOS HLA00690 - - - - - - Prof Antonio Arnaiz-Villena
DRB1*040501 - SW HLA00690 - - - - - - Dr KyuBum Kwack
DRB1*040502 DRB1*KOM KOM HLA00691 Moribe T, Kaneshige T, Hirakata M, Mimori T, Akitsuki M, Inoko H Identification of a DRB1*0405 variant (DRB1*04052) using the PCR-RFLP method Tissue Antigens 1996 47 450-3 Dr Toshihiko Kaneshige
DRB1*040503 DRB1*JVASA JVASA HLA01551 Greville WD, Chapman G, Hogbin JP, Dunckley H Novel HLA-DRB1 alleles revealed by sequencing based typing, DRB1*04053 and DRB1*1143 Tissue Antigens 2002 59 347-9 Dr Heather Dunckley
DRB1*040503 - 140383 HLA01551 - - - - - - Dr Heather Dunckley
DRB1*040504 - GN00419 HLA01605 Lazaro AM, Steiner NK, Moraes ME, Moraes JR, Ng J, Hartzman RJ, Hurley CK Ten novel HLA-DRB1 alleles and one novel DRB3 allele Tissue Antigens 2005 66 327-9 Dr Carolyn K Hurley
DRB1*040504 - HAN-1025 HLA01605 - - - - - - Ms Georgena Wohlwend
DRB1*040505 - BY00236 HLA03055 - - - - - - Dr Carolyn K Hurley
DRB1*040505 - R030242430 HLA03055 - - - - - - Ms Linda Smith
DRB1*040505 - BY00251 HLA03055 - - - - - - Dr Carolyn K Hurley
DRB1*040506 - JMDP01K028 HLA03375 - - - - - - Dr Kenji Tadokoro
DRB1*040601 KT2 LKT2 HLA00692 Gregersen PK, Goyert SM, Song QL, Silver J Microheterogeneity of HLA-DR4 haplotypes: DNA sequence analysis of LD"KT2" and LD"TAS" haplotypes Hum Immunol 1987 19 287-92 Dr Peter Gregersen
DRB1*040601 - 43A3 HLA00692 - - - - - - Prof Antonio Arnaiz-Villena
DRB1*040602 - CTM-9096471 HLA02172 Balas A, Vilches C, Rodriguez MA, Fernandez B, Martinez MP, de Pablo R, Garcia-Sanchez F, Vicario JL Group-specific amplification of cDNA from DRB1 genes. Complete coding sequences of partially defined alleles and identification of the new alleles DRB1*040602, DRB1*111102, DRB1*080103, and DRB1*0113. Human Immunology 2006 67 1008-16 Dr Antonio Balas
DRB1*040701 Dw13.2 R88 HLA00693 Lanchbury JS, Hall MA, Welsh KI, Panayi GS Sequence analysis of HLA-DR4B1 subtypes: additional first domain variability is detected by oligonucleotide hybridization and nucleotide sequencing Hum Immunol 1990 27 136-44 Dr Jerry S Lanchbury
DRB1*040701 - JHAF HLA00693 Lang B, Navarrete C, LoGalbo PR, Nepom GT, Silver J, Winchester RJ, Gregersen PK Further DNA sequence microheterogeneity of the HLA-DR4/Dw13 haplotype group: importance of amino acid position 86 of the DR beta 1 chain for T-cell recognition Hum Immunol 1990 27 378-89 Prof Bernhard Lang
DRB1*040701 - JRR HLA00693 - - - - - - Prof Antonio Arnaiz-Villena
DRB1*040701 - CTM-0096476 HLA00693 Balas A, Vilches C, Rodriguez MA, Fernandez B, Martinez MP, de Pablo R, Garcia-Sanchez F, Vicario JL Group-specific amplification of cDNA from DRB1 genes. Complete coding sequences of partially defined alleles and identification of the new alleles DRB1*040602, DRB1*111102, DRB1*080103, and DRB1*0113. Human Immunology 2006 67 1008-16 Dr Antonio Balas
DRB1*040702 DRB1*0407var NT0019 HLA01453 Chen DS, Tang TF, Pulyaeva H, Slack R, Tu B, Wagage D, Li L, Perlee L, Ng J, Hartzman RJ, Katovich Hurley C Relative HLA-DRB1*04 allele frequencies in five United States populations found in a hematopoietic stem cell volunteer donor registry and seven new DRB1*04 alleles Hum Immunol 2002 63 665-72 Dr Carolyn K Hurley
DRB1*040703 - CH112 HLA01706 - - - - - - Dr Mary Sue Leffell
DRB1*0408 Dw14.2 M36 HLA00694 Lanchbury JS, Hall MA, Welsh KI, Panayi GS Sequence analysis of HLA-DR4B1 subtypes: additional first domain variability is detected by oligonucleotide hybridization and nucleotide sequencing Hum Immunol 1990 27 136-44 Dr Jerry S Lanchbury
DRB1*0408 DR4-CETUS RA1 HLA00694 Gregersen PK, Todd JA, Erlich HA, Long E, Servenius B, Choi E, Kao HT, Lee JS First domain sequence diversity of DR and DQ subregion alleles Immunobiology of HLA. Vol. 1 1989 - 1027-31 Dr Henry Erlich
DRB1*0408 - SUDNA02547 HLA00694 - - - - - - Dr Carl F Grumet
DRB1*0408 - RGR HLA00694 - - - - - - Prof Antonio Arnaiz-Villena
DRB1*0408 - CTM-1096458 HLA00694 Balas A, Vilches C, Rodriguez MA, Fernandez B, Martinez MP, de Pablo R, Garcia-Sanchez F, Vicario JL Group-specific amplification of cDNA from DRB1 genes. Complete coding sequences of partially defined alleles and identification of the new alleles DRB1*040602, DRB1*111102, DRB1*080103, and DRB1*0113. Human Immunology 2006 67 1008-16 Dr Antonio Balas
DRB1*0409 - R80 HLA00695 Lanchbury JS, Jaeger EE, Welsh KI, Panayi GS Nucleotide sequence of a novel HLA-DRB4B1 allele, DRB1*0409 Immunogenetics 1991 33 210-2 Dr Jerry S Lanchbury
DRB1*0409 - LC284 HLA00695 Vilches C, Sepulveda S, Balas A, Solis R, Aviles MJ, Estefania E, Gomez-Lozano N, Vicario JL, Depablo R Complete coding sequences and haplotypic associations of HLA-B*0707, -B*1524, -B*4405, -B*4802, -DRB1*0409, -DRB1*0411, -DRB1*1115, -DRB1*1305, and the novel allele -DRB1*0709. Group-specific amplification of cDNA from DRB1 alleles associated to DRB3 and DRB4 Tissue Antigens 2005 65 529-38 Dr Carlos Vilches
DRB1*0409 - GN00057 HLA00695 - - - - - - Dr Carolyn K Hurley
DRB1*0409 - GN00057 HLA00695 - - - - - - Dr Carolyn K Hurley
DRB1*0410 DR4.CB CB HLA00696 Petersdorf EW, Smith AG, Mickelson EM, Martin PJ, Hansen JA Ten HLA-DR4 alleles defined by sequence polymorphisms within the DRB1 first domain Immunogenetics 1991 33 267-75 Prof Effie Petersdorf
DRB1*0410 - ABCC60 HLA00696 Gao X, Veale A, Serjeantson SW HLA class II diversity in Australian aborigines: unusual HLA-DRB1 alleles Immunogenetics 1992 36 333-7 Prof Susan W Serjeantson
DRB1*0410 - EGR HLA00696 - - - - - - Prof Antonio Arnaiz-Villena
DRB1*0410 STF22421 T.D.DNA#22421 HLA00696 - - - - - - Ms Betty Lo
DRB1*0410 - CTM-6097095 HLA00696 Balas A, Vilches C, Rodriguez MA, Fernandez B, Martinez MP, de Pablo R, Garcia-Sanchez F, Vicario JL Group-specific amplification of cDNA from DRB1 genes. Complete coding sequences of partially defined alleles and identification of the new alleles DRB1*040602, DRB1*111102, DRB1*080103, and DRB1*0113. Human Immunology 2006 67 1008-16 Dr Antonio Balas
DRB1*0411 DR4.EC EC HLA00697 Petersdorf EW, Smith AG, Mickelson EM, Martin PJ, Hansen JA Ten HLA-DR4 alleles defined by sequence polymorphisms within the DRB1 first domain Immunogenetics 1991 33 267-75 Prof Effie Petersdorf
DRB1*0411 - HV846 HLA00697 Gao X, Veale A, Serjeantson SW HLA class II diversity in Australian aborigines: unusual HLA-DRB1 alleles Immunogenetics 1992 36 333-7 Prof Susan W Serjeantson
DRB1*0411 - HAA HLA00697 - - - - - - Prof Peter Stastny
DRB1*0411 DR*04 JMJ HLA00697 - - - - - - Prof Antonio Arnaiz-Villena
DRB1*0411 - C211 HLA00697 Vilches C, Sepulveda S, Balas A, Solis R, Aviles MJ, Estefania E, Gomez-Lozano N, Vicario JL, Depablo R Complete coding sequences and haplotypic associations of HLA-B*0707, -B*1524, -B*4405, -B*4802, -DRB1*0409, -DRB1*0411, -DRB1*1115, -DRB1*1305, and the novel allele -DRB1*0709. Group-specific amplification of cDNA from DRB1 alleles associated to DRB3 and DRB4 Tissue Antigens 2005 65 529-38 Dr Carlos Vilches
DRB1*0412 DRB1*AB2 ABO1078 HLA00698 Gao X, Veale A, Serjeantson SW HLA class II diversity in Australian aborigines: unusual HLA-DRB1 alleles Immunogenetics 1992 36 333-7 Prof Susan W Serjeantson
DRB1*0412 - 132935 HLA00698 - - - - - - Dr Heather Dunckley
DRB1*0413 DRB1*LEV LEV HLA00699 Petersdorf EW, Smith AG, Martin PJ, Hansen JA HLA-DRB1 first domain sequence for a novel DR4 allele designated DRB1*0413 Tissue Antigens 1992 40 267-8 Prof Effie Petersdorf
DRB1*0414 DR4 Dw10.2 VK HLA00700 Pile KD, Willcox N, Bell JI, Wordsworth BP A novel HLA-DR4 allele (DRB1*0414) in a patient with myasthenia gravis Tissue Antigens 1992 40 264-6 Dr Paul Wordsworth
DRB1*0414 - NT00980 HLA00700 - - - - - - Dr Carolyn K Hurley
DRB1*0415 - NIC
HOU		 HLA00701 Tiercy JM, Gebuhrer L, Betuel H, Mach B, Jeannet M A new HLA-DR4 allele with a DR11 alpha-helix sequence Tissue Antigens 1993 41 97-101 Dr Jean-Marie Tiercy
DRB1*0415 - TER308 HLA00701 - - - - - - Prof Rainer Blasczyk
DRB1*0416 DR4-BELF BEL5GB HLA00702 Middleton D, Hughes DJ, Trainor F, Graham CA, Savage DA An HLA-DRB1*04 first domain sequence (DRB1*0416) which differs from HLA-DRB1*0401 at codon 59 Tissue Antigens 1994 43 44-6 Prof Derek Middleton
DRB1*0416 - BEL5GB HLA00702 - - - - - - Dr Carolyn K Hurley
DRB1*0416 - BEL5GB HLA00702 - - - - - - Prof Rainer Blasczyk
DRB1*0417 DRB1*04SAM TOB-0070 HLA00703 Zhang S, Fernandez-Vina M, Falco M, Cerna M, Raimondi E, Stastny P A novel HLA-DRB1 allele (DRB1*0417) in South American Indians Immunogenetics 1993 38 463 Prof Peter Stastny
DRB1*0417 - BY00255 HLA00703 - - - - - - Dr Carolyn K Hurley
DRB1*0418 DRB1*04.N AI7
AI8		 HLA00704 Mehra NK, Bouwens AG, Naipal A, Rajalingam R, Grubic Z, Taneja V, Tilanus MG, Giphart MJ Asian Indian HLA-DR2-, DR4-, and DR52-related DR-DQ genotypes analyzed by polymerase chain reaction based nonradioactive oligonucleotide typing. Unique haplotypes and a novel DR4 subtype Hum Immunol 1994 39 202-10 Prof Marcel GJ Tilanus
DRB1*0418 - 74DR HLA00704 - - - - - - Dr Marcelo Fernandez Vina
DRB1*0418 - GN0222 HLA00704 - - - - - - Dr Carolyn K Hurley
DRB1*0419 DR4FK FK HLA00705 Smith AG, Petersdorf EW, Mickelson E, Guthrie LA, Regen L, Martin PJ, Hansen JA HLA-DRB1 first domain sequences of two new DR11 alleles and one novel DR4 allele Tissue Antigens 1993 42 533-5 Prof Effie Petersdorf
DRB1*0419 - G.A.DNA#7881 HLA00705 - - - - - - Ms Betty Lo
DRB1*0420 DRB1*04MC AD-7863
BM29/92		 HLA00706 Cassidy S, Lester S, Humphreys I, Bennett G, Hetzel P, McCluskey J A new HLA-DRB1*04 allele: DRB1*0420 Tissue Antigens 1995 45 353-5 Prof James McCluskey
DRB1*0421 DR4New SMH HLA00707 Keller E, Yao Z, Volgger A, Lang B, Albert ED A novel variant of DR4 (DRB1*0421) identified in a patient with polychondritis Immunogenetics 1995 41 171 Prof Ekkehard D Albert
DRB1*0422 DR4New D18002 HLA00708 Varney MD, Kanaan C, Tait BD Identification of DRB1*0422--implications for serological epitopes Tissue Antigens 1996 47 150-2 Dr Michael Varney
DRB1*0423 DRB1*04New MAG HLA00709 Gebuhrer L, Adami N, Javaux F, Freidel AC, Jeannet M, Betuel H, Tiercy JM Sequence of a new HLA-DR4 allele with an unusual residue at position 88 that does not seem to affect T-cell allo recognition Hum Immunol 1996 51 60-2 Dr Jean-Marie Tiercy
DRB1*0424 DRB1*Mi M.i HLA00710 Dormoy A, Reviron DV, Froelich N, Weiller PJ, Mercier PJ, Tongio MM Birth of a new allele in a sibling: cis or trans gene conversion during meiosis? Immunogenetics 1997 46 520-3 Dr Anne Dormoy
DRB1*0425 DRB1*04ISA RI
HB		 HLA00711 Voorter CE, Hepkema BG, Lems SP, van den Berg-Loonen EM Identification of three new DRB1 alleles, DRB1*0107, *0425 and *13012 and confirmation of DRB4*01033 Tissue Antigens 2003 61 398-402 Prof Ella M van den Berg Loonen
DRB1*0426 DRB1*04CMT T010148 HLA00712 Buyse IM, Ouellet S, Hashemi-Tavoularis S Identification of novel DRB1*13 (DRB1*1333), DRB1*04 (DRB1*0426) and DRB5* (DRB5*0109) alleles Tissue Antigens 1998 51 658-62 Dr Sofia Tavoularis
DRB1*0426 - HAN-1024 HLA00712 - - - - - - Ms Georgena Wohlwend
DRB1*0427 DRB1*0403Variant NOR03 HLA00713 Chen DS, Tang TF, Pulyaeva H, Slack R, Tu B, Wagage D, Li L, Perlee L, Ng J, Hartzman RJ, Katovich Hurley C Relative HLA-DRB1*04 allele frequencies in five United States populations found in a hematopoietic stem cell volunteer donor registry and seven new DRB1*04 alleles Hum Immunol 2002 63 665-72 Dr Carolyn K Hurley
DRB1*0428 DRB1*0405V1 JC4772 HLA00714 - - - - - - Dr Kouichi Kashiwase
DRB1*0429 DRB1*0405V2 JC7616 HLA00715 - - - - - - Dr Kouichi Kashiwase
DRB1*0430 DRB1*0405V3 JC9227 HLA00716 - - - - - - Dr Kouichi Kashiwase
DRB1*0431 DRB1*04New GE47192 HLA00717 Bettens F, Kervaire B, Tiercy JM Sequence of HLA-DRB1*0431 allele Tissue Antigens 1999 53 378-80 Dr Jean-Marie Tiercy
DRB1*0431 - GE47192 HLA00717 - - - - - - Prof Rainer Blasczyk
DRB1*0432 DRB1*04-A NIE HLA00718 Anholts JD, Verduijn W, Drabbels J, Mulder A, Doxiadis II, Schreuder GM Identification of two new alleles HLA-DRB1*0312, DRB1*0432 and of a DRB3-negative DRB1*1313-positive haplotype Tissue Antigens 2000 56 87-9 Dr Willem Verduyn
DRB1*0433 DRB1*04_7468 WBD7468 HLA01088 - - - - - - Dr David Sayer
DRB1*0433 - 177927 HLA01088 - - - - - - Dr Heather Dunckley
DRB1*0434 DRB1*04new CB1653 HLA01167 - - - - - - Miss Kathleen Lebeer
DRB1*0434 - DWEVE488 HLA01167 - - - - - - Prof Rainer Blasczyk
DRB1*0435 DRB1*04New NT0009 HLA01235 Chen DS, Tang TF, Pulyaeva H, Slack R, Tu B, Wagage D, Li L, Perlee L, Ng J, Hartzman RJ, Katovich Hurley C Relative HLA-DRB1*04 allele frequencies in five United States populations found in a hematopoietic stem cell volunteer donor registry and seven new DRB1*04 alleles Hum Immunol 2002 63 665-72 Dr Carolyn K Hurley
DRB1*0436 - BN61 HLA01242 Chu CC, Lee HL, Lin M Identification of a new HLA-DRB1 allele, HLA-DRB1*0436 Tissue Antigens 2004 63 279-81 Dr Marie Lin
DRB1*0436 - 175586 HLA01242 - - - - - - Dr Heather Dunckley
DRB1*0437 DRB1*04nv MDPH0002764 HLA01338 Palou E, Campos E, Gil J, Santos M, Nogues N, Ribera A Identification of a new HLA-DRB1*04 allele, DRB1*0437 Tissue Antigens 2001 58 187-9 Dr Eduard Palou
DRB1*0438 - SLTA HLA01345 - - - - - - Dr Ryad Tamouza
DRB1*0438 - VTIS72428 HLA01345 - - - - - - Dr Michael Varney
DRB1*0439 DRB1*04031var NT0024 HLA01458 Chen DS, Tang TF, Pulyaeva H, Slack R, Tu B, Wagage D, Li L, Perlee L, Ng J, Hartzman RJ, Katovich Hurley C Relative HLA-DRB1*04 allele frequencies in five United States populations found in a hematopoietic stem cell volunteer donor registry and seven new DRB1*04 alleles Hum Immunol 2002 63 665-72 Dr Carolyn K Hurley
DRB1*0439 - 166438 HLA01458 - - - - - - Dr Heather Dunckley
DRB1*0440 DRB1*0404var NT0020 HLA01454 Chen DS, Tang TF, Pulyaeva H, Slack R, Tu B, Wagage D, Li L, Perlee L, Ng J, Hartzman RJ, Katovich Hurley C Relative HLA-DRB1*04 allele frequencies in five United States populations found in a hematopoietic stem cell volunteer donor registry and seven new DRB1*04 alleles Hum Immunol 2002 63 665-72 Dr Carolyn K Hurley
DRB1*0440 - 101330 HLA01454 - - - - - - Dr Heather Dunckley
DRB1*0440 - HAN-1023 HLA01454 - - - - - - Ms Georgena Wohlwend
DRB1*0441 DRB1*04031var NT0021 HLA01459 Chen DS, Tang TF, Pulyaeva H, Slack R, Tu B, Wagage D, Li L, Perlee L, Ng J, Hartzman RJ, Katovich Hurley C Relative HLA-DRB1*04 allele frequencies in five United States populations found in a hematopoietic stem cell volunteer donor registry and seven new DRB1*04 alleles Hum Immunol 2002 63 665-72 Dr Carolyn K Hurley
DRB1*0441 - NT00791 HLA01459 - - - - - - Dr Carolyn K Hurley
DRB1*0442 - JH71321 HLA01457 Dunn TA, Iglehart BA, Leffell MS Identification of a novel DR4 allele, DRB1*0442 Tissue Antigens 2001 58 198-200 Dr Mary Sue Leffell
DRB1*0443 - OORCH18 HLA01499 Chen DS, Tang TF, Pulyaeva H, Slack R, Tu B, Wagage D, Li L, Perlee L, Ng J, Hartzman RJ, Katovich Hurley C Relative HLA-DRB1*04 allele frequencies in five United States populations found in a hematopoietic stem cell volunteer donor registry and seven new DRB1*04 alleles Hum Immunol 2002 63 665-72 Dr Carolyn K Hurley
DRB1*0443 - OORCH18 HLA01499 - - - - - - Dr Jianming (James) Tang
DRB1*0444 - satt44124 HLA01601 - - - - - - Mr Ian Humphreys
DRB1*0445 - HY-LHB
HY-LBG		 HLA01695 Bae SC, Lee KW Identification of a novel DRB1*04 allele: DRB1*0445 Tissue Antigens 2003 62 270-2 Dr Kyung Wha Lee
DRB1*0446 - TBC-22381 HLA01746 - - - - - - Dr Masahiro Satake
DRB1*0447 - 119244 HLA01780 Velickovic ZM, Hogbin JP, Truong J, Jaafar S, Dunckley H Identification of two novel HLA class II alleles, DQB1*030503 and DRB1*0447 Tissue Antigens 2004 63 93-5 Dr Heather Dunckley
DRB1*0448 - TBC27636 HLA01793 - - - - - -